Try setting your router to UPnP. See instructions here:
http://www.panasonic.co.jp/pcc/produ...ys_wrt54g.html. Then set limewire to UPnP in Tools>Options>Advanced>Firewall Config, tick the UPnP button, click apply. Make sure the listen on port is the one you allowed access through your firewall.
Don't think the usb adapter WUSB54GC is affecting it.